Gramma was born in her native habitat and was estimated to be about 141 years old, zoo officials said. She died Nov. 20.

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Gramma, a Galápagos tortoise described by her care specialists as “the queen of the zoo,” was euthanized last Thursday after ...

Gramma the Galápagos tortoise dies at 141 at the San Diego Zoo ...

Waymo has announced that its driverless rideshare vehicles will be available to riders in San Diego in mid-2026.